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If the only available genes produce rather large changes, disadvantageous one at a time, then it seems to me probable that evolution will not occur in a random mating population. In a self-fertilised or highly inbred species it may do so if several mutations useful in conjunction, but separately harmful, occur simultaneously. Such an event is rare, but must happen reasonably often in wheat...

— J. B. S. Haldane The Causes of Evolution (1932)

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An inordinate fondness for beetles.

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The Causes of Evolution (1932)

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The Causes of Evolution (1932)

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